Blog 4 - Improving Plants

Blog 4 - Improving Plants

Is the pecan self-pollinating or cross-pollinating?

The pecan tree has both female and male organs, but they are not located together, which is called monecious flowering. (Smith, n.d.). The catkins (male flowers) and the pistil (female flower) receptiveness occur at different times. This is why the pecan tree requires cross-cultivation of a compatible cultivar. This is not entirely true, however. The exact times of the fertility of both organs is depended per pecan cultivar. While the Burkett cultivar is unable to self-pollinate, the Western cultivar does have the ability to do so. (Smith, n.d.). Studies do show that cross-pollination has a positive effect on the size and ripening of the pecan nuts. (Byford, 2005).

Logically, if a crop has only one reproductive organ it is not able to self-pollinate. And if it does, looking at the time-span of the receptiveness of each organ helps identifying its ability to self-pollinate or not. Mostly, plants that are able to self-pollinate are also able to cross-pollinate. Another insight is the amount of pollen grains. Self-pollinating crops do not require large amounts for they are simply located near to the pistil. Cross-pollinating crops have larger amounts of pollen grains, for they have to be carried by the wind, water or animals first in order to reach the pistil. (Diffen, n.d.). In the case of the pecan tree, the pollen grains are available in such abundance that they would be able to pollinate all pecans in the world. Another indication that defining the pecan tree as a cross-pollinator is the right-full classification for this crop. (Smith, n.d.).
                                    
Figure 1 Pecan nuts, different cultivars (Grauke & Thompson, n.d.).

Breeding program for the American Pecan 

Several breeding programs have taken place in the United States and more are expected to follow. Looking at one in particular we will go into details why the breeding program was necessary and what the characteristics are of the cultivar that has been created.
The University of Georgia started a breeding program of pecan trees in 1998. The University has started up new pecan breeding programs since then. The newest published evaluation on the breeding and cultivar program is one that was conducted from July 2010 till June 2015. (University of Georgia, 2015). The humid conditions of the Southern States of the United States, where the pecan production is located, are the source of growing fungi. These fungi are the source of scab infections. The main dilemma in breeding is to find a cultivar that is both fungi resistant and has the desirable nut quality traits. (See Figure 1). Commercial pecan plantings require 8-11 applications of fungicides in order to remain profitable. Profitability of a commercial plantation is also an issue that is addressed by breeding new cultivars also plays an important role in the innovation of pecan growing. (University of Georgia, 2015).
The breeding program that was conducted by the University of Georgia was done in a traditional way. The pollination was done in a controlled matter. The nuts were harvested, stratified for six weeks and then planted again. The cultivars were selected on tree productivity, nut quality, phenology and insect and disease resistance. (University of Georgia, 2015).

The results of the plant breeding program

The new cultivars have a good scab pressure which could result in as much as 80% of their offspring (progenies) being discarded for scab susceptibility. With their sufficient nut quality traits as well as fungi resistance using these cultivars could result in production cost savings. While over the years pecan nut prices have only increased with 4% in the period 1980-2015 the production costs of the nuts have risen over 154% in that same period. The savings could be approximately up to $115 per acre per year. The reduction of the costs will be on fungicide usage and labour. (University of Georgia, 2015). With the stress on low production costs in the pecan industry the need of innovations on this field are very much needed.

If you plant a crop of the same genotype in different regions in your country, will the phenotype be the same?

To be able to understand the sub issue we will look at the definition of both genotype and phenotype.

Genotype
The genetic constitution of an individual organism. (Oxford Dictonaries, n.d.).

Phenotype
The observable characteristics of an individual resulting from the interaction of its genotype with the environment. (Oxford Dictionaries, n.d.).

So the phenotype = genotype + environmental conditions. So, basically how the plant ends up developing. While a plant may share its genetical identity with another, the environmental conditions will eventually determine how the plant grows and develops. To give examples, its supply of sunshine, water and its rivalry with other plants. There are many more parameters that will determine how fast the plant will grow. Not only the genotype will influence nut quality and production capacity, but also the environmental conditions of each individual tree.

References

Byford, R. (2005, May). Growth and Development of Pecan Nuts. Retrieved from New Mexico State University: http://aces.nmsu.edu/pubs/_h/H618.pdf
Colorado State University. (2004, March). What Are Transgenic Plants? Retrieved from Colorado State University: http://www.cls.casa.colostate.edu/TransgenicCrops/what.html
Diffen. (n.d.). Cross Pollination vs. Self Pollination. Retrieved from Diffen: http://www.diffen.com/difference/Cross_Pollination_vs_Self_Pollination
Grauke, L., & Thompson, T. (n.d.). Pecan cultivars. Retrieved from Agricultural Research Service, U.S. Dept. of Agriculture: http://cgru.usda.gov/carya/pecans/03/yba1912.jpg
Oxford Dictionaries. (n.d.). Phenotype. Retrieved from Oxford Dictionaries: https://en.oxforddictionaries.com/definition/phenotype
Oxford Dictonaries. (n.d.). Genotype. Retrieved from Oxford Dictonaries: https://en.oxforddictionaries.com/definition/genotype
Smith, M. (n.d.). Botany. Retrieved from Oklahoma State University: http://pecan.okstate.edu/html/introduction/id.htm

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